Oceanic islands are often considered natural test tubes for evolution due to their isolation and limited number of colonizing species, leading to remarkable evolutionary radiations. Understanding how biodiversity differs between island systems and nearby mainland environments has intrigued researchers for decades. The goal of the present study is to examine the influence of evolutionary constraints and ecological drivers of ecomorphology among land snail species forming assemblages found on an island system (the Galápagos Islands) and on a nearby mainland system (the Peruvian Andes). Land snails of the Family Bulimulidae were sampled across the two landscapes. Ecological and shell morphological data will be assembled and curated for all species encountered at each site. Evolutionary relationships among species will be inferred from genomic markers, and a phylogenetic framework will be used to estimate diversification rates and assess evolutionary constraints for each group. We will then use a multivariate analysis approach to examine the relationships and differences between shell morphology, using geometric morphometrics and ecological data at the species and community levels.